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Further, a more complete picture of the genetic and environmental aspects that impact com Robert Wright. Both nurture environmental and nature genetic factors play an important role in human disease etiology. Traditionally, these effects have been thought of as independent.

This perspective is ill informed for non-mendelian complex disorders which result as an interaction between genetics and environment. To understand health and disease we must study how nature and nurture interact. Recent advances in human genomics and high-throughput biotechnology make it possible to study large numbers of genetic markers and gene products simultaneously to explore their interactions with environment.

We present an expanded environmental genomic disease paradigm. We discuss several study design issues for gene-environmental interaction studies, including confounding and selection bias, measurement of e Xavier Estivill.